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When the Persian King Darius is slain during his charge to conquer Greece, a far greater threat is born. Fueled by revenge and manipulated by his rancorous commander Artemisia (Eva Green), Darius' son Xerxes (Rodrigo Santoro) transforms himself into the "god king" and rallies Persia's titanic army in a new campaign to destroy the city-states. Though vastly outnumbered, Athenian admiral Themistocles (Sullivan Stapleton) uses superior military strategies to triumph over the initial nautical onslaught from the Persians, but he soon realizes that without the aid of Sparta's elite warriors, all of Greece may fall.